Pinus palustris Mill.

Range and Habitat

The native range of longleaf pine. (From Little, 1971.)

Geographic Range


According to Boyer (1990) : "The natural range of longleaf pine includes most of the Atlantic and Gulf Coastal Plains from southeastern Virginia to eastern Texas and south through the northern two-thirds of peninsular Florida. The species also grows in the Piedmont, Ridge and Valley, and Mountain Provinces of Alabama and northwest Georgia."

Climate


According to Boyer (1990) : "Longleaf pine grows in warm, wet temperate climates characterized by hot summers and mild winters. Annual mean temperatures range from 16° to 23° C (60° to 74° F) and annual precipitation from 1090 to 1750 mm (43-50 in) in the Carolinas and Texas and the greatest along the Gulf Coast of Alabama, Mississippi, and extreme west Florida. A distinct summer rainfall peak occurs along the Atlantic Coast, being most pronounced in Florida. A secondary rainfall peak in March becomes pronounced along the Gulf Coast. Fall is the driest season of the year, although droughts during the growing season are not unusual."

Soils and Topography


According to Boyer (1990) : "Longleaf pine is native to a wide variety of sites ranging from wet, poorly drained flatwoods to dry, rocky mountain ridges. Elevations range from barely above sea level near the beaches on the lower Coastal Plain up to about 600 m (1970 ft) in the mountains of Alabama. Most of the longleaf pine forests are found on the Atlantic and Gulf Coastal plains at elevations below 200 m (660 ft). Here the soils are largely derived from marine sediments and range from deep coarse, excessively drained sands to poorly drained clays. For the most part, surface soils are sandy, acid, low in organic matter, and relatively infertile."

Three soil orders are associated with longleaf pine. The first, Ultisols, are the most expansive in the southeast, excluding peninsular Florida. Entisols make up the Sandhills of the southeastern Atlantic states. Spodosols, which are wet sandy soils, have a fluctuating water table near the ground surface during rainy seasons. This soil is common to flatwood sites of the lower Coastal Plain of Florida.

Forest Associates


According to Boyer (1990) : "The principal longleaf cover types are Longleaf Pine (Society of American Foresters Type 70), Longleaf Pine-Scrub Oak (Type 71), and Longleaf Pine-Slash Pine (Type 83). Longleaf pine is also a minor component of other forest types within its range: Sand Pine (Type 69), Shortleaf Pine (Type 75), Loblolly Pine (Type 81), Loblolly Pine-Hardwoods (Type 82), Slash Pine (Type 84), and South Florida Slash Pine (Type 111).

"Longleaf pine develops in close association with periodic surface fires. The vegetation associated with longleaf pine reflects the frequency and severity of burning. In the past, frequent fires resulted in open, parklike stands of longleaf with few other woody plants and a ground cover dominated by grasses. Ground cover in longleaf pine in the Coastal Plains can be separated into two general regions, with the division in the central part of south Alabama and northwest Florida. To the west, bluestem (Andropogon spp.) and panicum (Panicum spp.) grasses predominate; to the east, wiregrass (pineland threeawn, Aristida stricta) is most common.

"With a reduction in fire occurrence, hardwoods and other pines encroach on the longleaf forest. Within the range of slash pine (Pinus elliottii), this species becomes increasingly important, leading to the cover type Longleaf Pine-Slash Pine. Elsewhere loblolly and shortleaf pines (P. taeda and P. echinata as well as hardwoods gradually replace the longleaf, eventually resulting in Loblolly Pine-Hardwood (Type 82) or occasionally Loblolly Pine-Shortleaf (Type 80). On poor, dry sandhillls and mountain ridges, scrub hardwoods invade the understory creating forest cover type Longleaf Pine-Scrub Oak and finally Southern Scrub Oak (Type 72) as the pine disappears.

"Hardwoods most closely associated with longleaf pine on mesic Coastal Plain sites include southern red, blackjack, and water oaks (Quercus falcata, Q. marilandica, and Q. nigra); flowering dogwood (Cornus florida); blackgum (Nyssa sylvatica); sweetgum (Liquidambar styraciflua); persimmon (Diospyros virginiana); and sassafras (Sassafras albidum). The more common shrubs include gallberry (Ilex glabra), yaupon (I. vomitoria), southern bayberry (Myrica cerifera), shining sumac (Rhus copallina), blueberry (Vaccinium spp.), huckleberry (Gaylussacia spp.), and blackberry (Rubus spp.). On xeric sandhill sites, the most common associates are turkey, bluejack, blackjack, sand post, and dwarf live oaks (Quercus laevis, Q. incana, Q. marilandica, Q. stellata var. margaretta, and Q. minima). On the dry clay hills and mountains of Alabama, blackjack, post (Q. stellata), and southern red oaks, and mockernut hickory (Carya tomentosa) are found with longleaf pine. On low, wet flatwood sites near the coast, the most conspicuous understory plants are gallberry and saw-palmetto (Serenoa repens). Other common understory plants in low, wet Longleaf Pine or Longleaf Pine-Slash Pine types are sweetbay (Magnolia virginiana), swamp cyrilla (Ilex coriacea), buckwheat-tree (Cliftonia monophylla), blueberries, and blackberries."

Habitat Competition


According to Boyer (1990) : "Longleaf pine is intolerant of competition, whether for light or for moisture and nutrients. The species will grow best in the complete absence of all competition, including that from other memebers of the species. Fortunately, as noted earlier, young even-age longleaf pine stands break up rapidly into a broad range of size classes, due to variability in duration of the grass stage. Stagnation is almost never a problem. However, even suppressed trees in a stand will slow the growth of dominant neighbors. Optimum stand density for development of crop trees needs to be maintained by periodic thinning. Given release from neighboring trees, dominant and codominate trees in an over-dense stand will respond promptly with increased diameter growth, as will some intermediate trees that retain crown ratios of 30 percent or more. Suppressed trees, while they may continue to live, rarely respond to release with improved growth."

Hybrids


According to Boyer (1990) : "The major southern pines, as well as some minor species, are closely related and have overlapping ranges. Natural hybridization has contribute to genetic diversity among trees and populations. Natural hybridization is common between longleaf and loblolly pine, producing the Sonderegger pine (Pinus × sondereggeri H.H. Chapm.). This is the only named southern pine hybrid. Throughout much of the longleaf pine range, the flowering of longleaf and loblolly pines overlaps in most years so there is no phenological barrier to natural corssing. Natural hybridization between longleaf and slah pine is unlikely, based on differences between the species in dormancy and heat requirements for flowering.

"Artificial crosses between longleaf pine and both loblolly and slash pines can be achieved easily. Crosses between longleaf and shortleaf pne have not been found in nature but have been produced artificially. There are no reported successful crosses of longleaf pine with any other pine species."


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